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The Genetic CodesCompiled by Andrzej (Anjay) Elzanowski and Jim Ostell at National Center for Biotechnology Information (NCBI), Bethesda, Maryland, U.S.A. Last update of the Genetic Codes: Sep. 23, 2024NCBI takes great care to ensure that the translation for each coding sequence (CDS) present in GenBank records is correct. Central to this effort is careful checking on the taxonomy of each record and assignment of the correct genetic code (shown as a /transl_table qualifier on the CDS in the flat files) for each organism and record. This page summarizes and references this work. The synopsis presented below is based primarily on the reviews by Osawa et al. (1992) and Jukes and Osawa (1993). Listed in square brackets [] (under Systematic Range) are tentative assignments of a particular code based on sequence homology and/or phylogenetic relationships. The print-form ASN.1 version of this document, which includes all the genetic codes outlined below, is also available here. Detailed information on codon usage can be found at the Codon Usage Database. GenBank format by historical convention displays mRNA sequences using the DNA alphabet. Thus, for the convenience of people reading GenBank records, the genetic code tables shown here use T instead of U. The initiator codon - whether it is AUG, CTG, TTG or something else, - is by default translated as methionine (Met, M). The possible intiator codons are marked as 'M' in the second ('Starts') row of the translation tables. Currently, genetic codes can be set independently for nucleus, mitochondria, plastids and hydrogenosomes. The current settings for each of these on the taxonomic tree can be viewed by the four buttons directly underneath the following code list. The following genetic codes are described here:
1. The Standard Code (transl_table=1)By default all transl_table in GenBank flatfiles are equal to id 1, and this is not shown. When transl_table is not equal to id 1, it is shown as a qualifier on the CDS feature. TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA * Ter TGA * Ter TTG L Leu i TCG S Ser TAG * Ter TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu i CCG P Pro CAG Q Gln CGG R Arg ATT I Ile ACT T Thr AAT N Asn AGT S Ser ATC I Ile ACC T Thr AAC N Asn AGC S Ser ATA I Ile ACA T Thr AAA K Lys AGA R Arg ATG M Met i ACG T Thr AAG K Lys AGG R Arg GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val GCG A Ala GAG E Glu GGG G Gly Click here to change format Initiation Codon:AUG Alternative Initiation Codons:In rare cases, translation in eukaryotes can be initiated from codons other than AUG. A well documented case (including direct protein sequencing) is the GUG start of a ribosomal P protein of the fungus Candida albicans (Abramczyk et al.) and the GUG initiation in mammalian NAT1 (Takahashi et al. 2005). Other examples can be found in the following references: Peabody 1989; Prats et al. 1989; Hann et al. 1992; Sugihara et al. 1990. The standard code currently allows initiation from UUG and CUG in addition to AUG.Back to top 2. The Vertebrate Mitochondrial Code (transl_table=2)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA * Ter TGA W Trp TTG L Leu TCG S Ser TAG * Ter TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu CCG P Pro CAG Q Gln CGG R Arg ATT I Ile i ACT T Thr AAT N Asn AGT S Ser ATC I Ile i ACC T Thr AAC N Asn AGC S Ser ATA M Met i ACA T Thr AAA K Lys AGA * Ter ATG M Met i ACG T Thr AAG K Lys AGG * Ter GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val i GCG A Ala GAG E Glu GGG G Gly Click here to change format Differences from the Standard Code:
Code 2 Standard
AGA Ter * Arg R
AGG Ter * Arg R
AUA Met M Ile I
UGA Trp W Ter *
Alternative Initiation Codons:Bos: AUA Homo: AUA, AUU Mus: AUA, AUU, AUC Coturnix, Gallus: also GUG (Desjardins and Morais, 1991) Systematic Range:Vertebrata Comments:AGA and AGG were thought to have become mitochondrial stop codons early in vertebrate evolution (Osawa, Ohama, Jukes & Watanabe 1989). However, at least in humans it has now been shown that AGA and AGG sequences are not recognized as termination codons. A -1 mitoribosome frameshift occurs at the AGA and AGG codons predicted to terminate the CO1 and ND6 ORFs, and consequently both ORFs terminate in the standard UAG codon (Temperley et al. 2010). Mitochondrial genes in some vertebrate (including humans) have incomplete stop codons ending in U or UA, which become complete termination codons (UAA) upon subsequent polyadenylation (Hou et al. 2006; Oh et al. 2007; Ki et al. 2010; Temperley R J et al 2010).Back to top 3. The Yeast Mitochondrial Code (transl_table=3)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA * Ter TGA W Trp TTG L Leu TCG S Ser TAG * Ter TGG W Trp CTT T Thr CCT P Pro CAT H His CGT R Arg CTC T Thr CCC P Pro CAC H His CGC R Arg CTA T Thr CCA P Pro CAA Q Gln CGA R Arg CTG T Thr CCG P Pro CAG Q Gln CGG R Arg ATT I Ile ACT T Thr AAT N Asn AGT S Ser ATC I Ile ACC T Thr AAC N Asn AGC S Ser ATA M Met i ACA T Thr AAA K Lys AGA R Arg ATG M Met i ACG T Thr AAG K Lys AGG R Arg GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val i GCG A Ala GAG E Glu GGG G Gly Click here to change format Differences from the Standard Code:
Code 3 Standard
AUA Met M Ile I
CUU Thr T Leu L
CUC Thr T Leu L
CUA Thr T Leu L
CUG Thr T Leu L
UGA Trp W Ter *
Systematic Range:Saccharomyces cerevisiae, Candida glabrata, Hansenula saturnus, and Kluyveromyces thermotolerans (Clark-Walker and Weiller, 1994) Comments:GUG (GTG) is used as a start codon for a few proteins in some Saccharomyces species (Sulo et al. 2017). The remaining CGN codons are rare in Saccharomyces cerevisiae and absent in Candida glabrata (= Torulopsis glabrata). The AUA codon is common in the gene var1 coding for the single mitochondrial ribosomal protein, but rare in genes encoding the enzymes. The coding assignments of the AUA (Met or Ile) and CUU (possibly Leu, not Thr) are uncertain in Hansenula saturnus. The coding assignment of Thr to CUN is uncertain in Kluyveromyces thermotolerans (Clark-Walker and Weiller, 1994).Back to top 4. The Mold, Protozoan, and Coelenterate Mitochondrial Code and the Mycoplasma/Spiroplasma Code (transl_table=4)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu i TCA S Ser TAA * Ter TGA W Trp TTG L Leu i TCG S Ser TAG * Ter TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu i CCG P Pro CAG Q Gln CGG R Arg ATT I Ile i ACT T Thr AAT N Asn AGT S Ser ATC I Ile i ACC T Thr AAC N Asn AGC S Ser ATA I Ile i ACA T Thr AAA K Lys AGA R Arg ATG M Met i ACG T Thr AAG K Lys AGG R Arg GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val i GCG A Ala GAG E Glu GGG G Gly Click here to change format Differences from the Standard Code:
Code 4 Standard
UGA Trp W Ter *
Alternative Initiation Codons:Trypanosoma: UUA, UUG, CUG Leishmania: AUU, AUA Tertrahymena: AUU, AUA, AUG Paramecium: AUU, AUA, AUG, AUC, GUG, GUA(?) Systematic Range:Bacteria: The code is used in Entomoplasmatales and Mycoplasmatales (Bove et al. 1989). The situation in the Acholeplasmatales is unclear. Based on a study of ribosomal protein genes, it had been concluded that UGA does not code for tryptophan in plant-pathogenic mycoplasma-like organisms (MLO) and the Acholeplasmataceae (Lim and Sears, 1992) and there seems to be only a single tRNA-CCA for tryptophan in Acholeplasma laidlawii (Tanaka et al. 1989). In contrast, in a study of codon usage in Phytoplasmas, it was found that 30 out of 78 ORFs analyzed translated better with code 4 (UGA for tryptophan) than with code 11 while the remainder showed no differences between the two codes (Melamed et al. 2003). In addition, the coding reassignment of UGA Stop --> Trp can be found in an alpha-proteobacterial symbiont of cicadas: Candidatus Hodgkinia cicadicola (McCutcheon et al. 2009). Fungi: Emericella nidulans, Neurospora crassa, Podospora anserina, Acremonium (Fox, 1987), Candida parapsilosis (Guelin et al., 1991), Trichophyton rubrum (de Bievre and Dujon, 1992), Dekkera/Brettanomyces, Eeniella (Hoeben et al., 1993), and probably Ascobolus immersus, Aspergillus amstelodami, Claviceps purpurea, and Cochliobolus heterostrophus. Other Eukaryotes: Gigartinales among the red algae (Boyen et al. 1994), and the protozoa Trypanosoma brucei, Leishmania tarentolae, Paramecium tetraurelia, Tetrahymena pyriformis and probably Plasmodium gallinaceum (Aldritt et al., 1989). Metazoa: Coelenterata (Ctenophora and Cnidaria) Comments:This code is also used for the kinetoplast DNA (maxicircles, minicircles). Kinetoplasts are modified mitochondria (or their parts).Back to top 5. The Invertebrate Mitochondrial Code (transl_table=5)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA * Ter TGA W Trp TTG L Leu i TCG S Ser TAG * Ter TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu CCG P Pro CAG Q Gln CGG R Arg ATT I Ile i ACT T Thr AAT N Asn AGT S Ser ATC I Ile i ACC T Thr AAC N Asn AGC S Ser ATA M Met i ACA T Thr AAA K Lys AGA S Ser ATG M Met i ACG T Thr AAG K Lys AGG S Ser GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val i GCG A Ala GAG E Glu GGG G Gly Click here to change format Comment:The codon AGG is absent in Drosophila. Differences from the Standard Code:
Code 5 Standard
AGA Ser S Arg R
AGG Ser S Arg R
AUA Met M Ile I
UGA Trp W Ter *
Alternative Initiation Codons:AUA, AUU AUC: Apis (Crozier and Crozier, 1993) GUG: Polyplacophora (Boore and Brown, 1994 GenBank Accession Number: UUG: Ascaris, Caenorhabditis Systematic Range:Nematoda: Ascaris, Caenorhabditis; Mollusca: Bivalvia (Hoffmann et al., 1992); Polyplacophora (Boore and Brown, 1994) Arthropoda/Crustacea: Artemia (Batuecas et al., 1988); Arthropoda/Insecta: Drosophila [Locusta migratoria (migratory locust), Apis mellifera (honeybee)] Comments:Several arthropods translate the codon AGG as lysine instead of serine (as in the invertebrate mitochondrial genetic code) or arginine (as in the standard genetic code) (Abascal et al., 2006). GUG may possibly function as an initiator in Drosophila (Clary and Wolstenholme, 1985; Gadaleta et al., 1988). AUU is not used as an initiator in Mytilus (Hoffmann et al., 1992). "An exceptional mechanism must operate for initiation of translation of the cytochrome oxidase subunit I mRNA in both D. melanogaster (de Bruijn, 1983) and D. yakuba (Clary and Wolstenholme 1983), since its only plausible initiation codon, AUA, is out of frame with the rest of the gene. Initiation appears to require the "reading" of of an AUAA quadruplet, which would be equivalent to initiation at AUA followed immediately by a specific ribosomal frameshift. Another possible mechanism ... is that the mRNA is "edited" to bring the AUA initiation into frame." (Fox, 1987)Back to top 6. The Ciliate, Dasycladacean and Hexamita Nuclear Code (transl_table=6)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA Q Gln TGA * Ter TTG L Leu TCG S Ser TAG Q Gln TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu CCG P Pro CAG Q Gln CGG R Arg ATT I Ile ACT T Thr AAT N Asn AGT S Ser ATC I Ile ACC T Thr AAC N Asn AGC S Ser ATA I Ile ACA T Thr AAA K Lys AGA R Arg ATG M Met i ACG T Thr AAG K Lys AGG R Arg GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val GCG A Ala GAG E Glu GGG G Gly Click here to change format Differences from the Standard Code:
Code 6 Standard
UAA Gln Q Ter *
UAG Gln Q Ter *
Systematic Range:Ciliata: Oxytricha and Stylonychia (Hoffman et al. 1995), Paramecium, Tetrahymena, Oxytrichidae and probably Glaucoma chattoni. Dasycladaceae: Acetabularia (Schneider et al., 1989) and Batophora (Schneider and de Groot, 1991). Diplomonadida: Scope: Hexamita inflata, Diplomonadida ATCC50330, and ATCC50380. Ref.: Keeling, P.J. and Doolittle, W.F. 1996.. A non-canonical genetic code in an early diverging eukaryotic lineage. The EMBO Journal 15, 2285-2290. Comment:The ciliate macronuclear code has not been determined completely. The codon UAA is known to code for Gln only in the Oxytrichidae.Back to top 9. The Echinoderm and Flatworm Mitochondrial Code (transl_table=9)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA * Ter TGA W Trp TTG L Leu TCG S Ser TAG * Ter TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu CCG P Pro CAG Q Gln CGG R Arg ATT I Ile ACT T Thr AAT N Asn AGT S Ser ATC I Ile ACC T Thr AAC N Asn AGC S Ser ATA I Ile ACA T Thr AAA N Asn AGA S Ser ATG M Met i ACG T Thr AAG K Lys AGG S Ser GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val i GCG A Ala GAG E Glu GGG G Gly Click here to change format Differences from the Standard Code:
Code 9 Standard
AAA Asn N Lys K
AGA Ser S Arg R
AGG Ser S Arg R
UGA Trp W Ter *
Systematic Range:Asterozoa (starfishes) (Himeno et al., 1987) Echinozoa (sea urchins) (Jacobs et al., 1988; Cantatore et al., 1989) Rhabditophora among the Platyhelminthes (Telford et al. 2000)Back to top 10. The Euplotid Nuclear Code (transl_table=10)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA * Ter TGA C Cys TTG L Leu TCG S Ser TAG * Ter TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu CCG P Pro CAG Q Gln CGG R Arg ATT I Ile ACT T Thr AAT N Asn AGT S Ser ATC I Ile ACC T Thr AAC N Asn AGC S Ser ATA I Ile ACA T Thr AAA K Lys AGA R Arg ATG M Met i ACG T Thr AAG K Lys AGG R Arg GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val GCG A Ala GAG E Glu GGG G Gly Click here to change format Differences from the Standard Code:
Code 10 Standard
UGA Cys C Ter *
Systematic Range:Ciliata: Euplotidae (Hoffman et al. 1995).Back to top 11. The Bacterial, Archaeal and Plant Plastid Code (transl_table=11)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA * Ter TGA * Ter TTG L Leu i TCG S Ser TAG * Ter TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu i CCG P Pro CAG Q Gln CGG R Arg ATT I Ile i ACT T Thr AAT N Asn AGT S Ser ATC I Ile i ACC T Thr AAC N Asn AGC S Ser ATA I Ile i ACA T Thr AAA K Lys AGA R Arg ATG M Met i ACG T Thr AAG K Lys AGG R Arg GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val i GCG A Ala GAG E Glu GGG G Gly Click here to change format Systematic Range and Comments:Table 11 is used for Bacteria, Archaea, prokaryotic viruses and chloroplast proteins. As in the standard code, initiation is most efficient at AUG. In addition, GUG and UUG starts are documented in Archaea and Bacteria (Kozak 1983, Fotheringham et al. 1986, Golderer et al. 1995, Nolling et al. 1995, Sazuka & Ohara 1996, Genser et al. 1998, Wang et al. 2003). In E. coli, UUG is estimated to serve as initiator for about 3% of the bacterium's proteins (Blattner et al. 1997). CUG is known to function as an initiator for one plasmid-encoded protein (RepA) in Escherichia coli (Spiers and Bergquist, 1992). In addition to the NUG initiations, in rare cases Bacteria can initiate translation from an AUU codon as e.g. in the case of poly(A) polymerase PcnB and the InfC gene that codes for translation initiation factor IF3 (Polard et al. 1991, Liveris et al. 1993, Sazuka & Ohara 1996, Binns & Masters 2002). The internal assignments are the same as in the standard code though UGA codes at low efficiency for Trp in Bacillus subtilis and, presumably, in Escherichia coli (Hatfiled and Diamond, 1993).Back to top 12. The Alternative Yeast Nuclear Code (transl_table=12)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA * Ter TGA * Ter TTG L Leu TCG S Ser TAG * Ter TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG S Ser i CCG P Pro CAG Q Gln CGG R Arg ATT I Ile ACT T Thr AAT N Asn AGT S Ser ATC I Ile ACC T Thr AAC N Asn AGC S Ser ATA I Ile ACA T Thr AAA K Lys AGA R Arg ATG M Met i ACG T Thr AAG K Lys AGG R Arg GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val GCG A Ala GAG E Glu GGG G Gly Click here to change format Differences from the Standard Code:
Code 12 Standard
CUG Ser Leu
Alternative Initiation Codons:CAG may be used in Candida albicans (Santos et al., 1993). Systematic Range and Comments:In code 12 the CUG codon has been reassigned from Leu to Ser (Santos et al., 2011, Mateus et al., 2013, Krassowski et al., 2018). This code is used by the fungal CUG-Ser1 and CUG-Ser2 clades that represent separate monophyletic lineages of the Saccharomycotina (Li et al., 2021). In the NCBI taxonomy, CUG-Ser1 currently comprises the three families Cephaloascaceae, Debaryomycetaceae and Metschnikowiaceae and the genus Babjeviella. The CUG-Ser1 clade includes important human pathogens like Candida albicans and the multidrug-resistant Candida auris. The CUG-Ser2 clade represents a parallel reassignment of CUG and includes species in the families Ascoideaceae and Saccharomycopsidaceae. Most other true yeasts (Saccharomycotina) use the standard code however.Back to top 13. The Ascidian Mitochondrial Code (transl_table=13)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA * Ter TGA W Trp TTG L Leu i TCG S Ser TAG * Ter TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu CCG P Pro CAG Q Gln CGG R Arg ATT I Ile ACT T Thr AAT N Asn AGT S Ser ATC I Ile ACC T Thr AAC N Asn AGC S Ser ATA M Met i ACA T Thr AAA K Lys AGA G Gly ATG M Met i ACG T Thr AAG K Lys AGG G Gly GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val i GCG A Ala GAG E Glu GGG G Gly Click here to change format Differences from the Standard Code:
Code 13 Standard
AGA Gly G Arg R
AGG Gly G Arg R
AUA Met M Ile I
UGA Trp W Ter *
Systematic range and Comments:There is evidence from a phylogenetically diverse sample of tunicates (Urochordata) that AGA and AGG code for glycine. In other organisms, AGA/AGG code for either arginine or serine and in vertebrate mitochondria they code a STOP. Evidence for glycine translation of AGA/AGG has been found in Pyura stolonifera (Durrheim et al. 1993), Halocynthia roretzi (Kondow et al. 1999,Yokobori et al., 1993, Yokobori et al. 1999) and Ciona savignyi (Yokobori et al. 2003). In addition, the Halocynthia roretzi mitochondrial genome encodes an additional tRNA gene with the anticodon U*CU that is thought to enable the use of AGA or AGG codons for glycine and the gene has been shown to be transcribed in vivo (Kondow et al. 1999, Yokobori et al. 1999). Alternative initiation codons:ATA, GTG and TTG (Yokobori et al. 1999). ATT is the start codon for the CytB gene in Halocynthia roretzi (Gissi and Pesole, 2003).Back to top 14. The Alternative Flatworm Mitochondrial Code (transl_table=14)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA Y Tyr TGA W Trp TTG L Leu TCG S Ser TAG * Ter TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu CCG P Pro CAG Q Gln CGG R Arg ATT I Ile ACT T Thr AAT N Asn AGT S Ser ATC I Ile ACC T Thr AAC N Asn AGC S Ser ATA I Ile ACA T Thr AAA N Asn AGA S Ser ATG M Met i ACG T Thr AAG K Lys AGG S Ser GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val GCG A Ala GAG E Glu GGG G Gly Click here to change format Differences from the Standard Code:
Code 14 Standard
AAA Asn N Lys K
AGA Ser S Arg R
AGG Ser S Arg R
UAA Tyr Y Ter *
UGA Trp W Ter *
Systematic Range:Platyhelminthes (flatworms) and Nematoda (roundworms) Comments:Code 14 differs from code 9 only by translating UAA to Tyr rather than STOP. A recent study (Telford et al. 2000) has found no evidence that the codon UAA codes for Tyr in the flatworms but other opinions exist. There are very few GenBank records that are translated with code 14 but a test translation shows that retranslating these records with code 9 can cause premature terminations. More recently, UAA has been found to code for tyrosine in the nematodes Radopholus similis and Radopholus arabocoffeae (Jacob et al. 2009).Back to top 15. Blepharisma Nuclear Code (transl_table=15)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA * Ter TGA * Ter TTG L Leu TCG S Ser TAG Q Gln TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu CCG P Pro CAG Q Gln CGG R Arg ATT I Ile ACT T Thr AAT N Asn AGT S Ser ATC I Ile ACC T Thr AAC N Asn AGC S Ser ATA I Ile ACA T Thr AAA K Lys AGA R Arg ATG M Met i ACG T Thr AAG K Lys AGG R Arg GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val GCG A Ala GAG E Glu GGG G Gly Click here to change format Differences from the Standard Code:
Code 15 Standard
TAG Gln Q STOP
Systematic Range:Crassvirales: Experimental validation that human microbiome phages use alternative genetic coding. Peters et al., Nature Communications 2022Back to top 16. Chlorophycean Mitochondrial Code (transl_table=16)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA * Ter TGA * Ter TTG L Leu TCG S Ser TAG L Leu TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu CCG P Pro CAG Q Gln CGG R Arg ATT I Ile ACT T Thr AAT N Asn AGT S Ser ATC I Ile ACC T Thr AAC N Asn AGC S Ser ATA I Ile ACA T Thr AAA K Lys AGA R Arg ATG M Met i ACG T Thr AAG K Lys AGG R Arg GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val GCG A Ala GAG E Glu GGG G Gly Click here to change format Differences from the Standard Code:
Code 16 Standard
TAG Leu L STOP
Systematic Range:Chlorophyceae (Hayashi-Ishimaru et al. 1996. UAG is a sense codon in several chlorophycean mitochondria) and the chytridiomycete fungus Spizellomyces punctatus (Laforest et al. 1997. Mitochondrial tRNAs in the lower fungus Spizellomyces punctatus: tRNA editing and UAG 'stop' codons recognized as leucine).Back to top 21. Trematode Mitochondrial Code (transl_table=21)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA * Ter TGA W Trp TTG L Leu TCG S Ser TAG * Ter TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu CCG P Pro CAG Q Gln CGG R Arg ATT I Ile ACT T Thr AAT N Asn AGT S Ser ATC I Ile ACC T Thr AAC N Asn AGC S Ser ATA M Met ACA T Thr AAA N Asn AGA S Ser ATG M Met i ACG T Thr AAG K Lys AGG S Ser GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val i GCG A Ala GAG E Glu GGG G Gly Click here to change format Differences from the Standard Code:
Code 21 Standard
TGA Trp W STOP
ATA Met M Ile
AGA Ser S Arg
AGG Ser S Arg
AAA Asn N Lys
Systematic Range:Trematoda: Ohama, T, S. Osawa, K. Watanabe, T.H. Jukes, 1990. J. Molec Evol. 30 Garey, J.R. and D.R. Wolstenholme, 1989. J. Molec. Evol. 28: 374-387 329-332.Back to top 22. Scenedesmus obliquus Mitochondrial Code (transl_table=22)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA * Ter TAA * Ter TGA * Ter TTG L Leu TCG S Ser TAG L Leu TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu CCG P Pro CAG Q Gln CGG R Arg ATT I Ile ACT T Thr AAT N Asn AGT S Ser ATC I Ile ACC T Thr AAC N Asn AGC S Ser ATA I Ile ACA T Thr AAA K Lys AGA R Arg ATG M Met i ACG T Thr AAG K Lys AGG R Arg GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val GCG A Ala GAG E Glu GGG G Gly Click here to change format Differences from the Standard Code:
Code 22 Standard
TCA STOP * Ser
TAG Leu L STOP
Systematic Range:Scenedesmus obliquus: Nedelcu A, Lee RW, Lemieux C, Gray MW and Burger G. "The complete mitochondrial DNA sequence of Scenedesmus obliquus reflects an intermediate stage in the evolution of the green algal mitochondrial genome." Genome Res. 2000 Jun;10(6):819-31.Back to top 23. Thraustochytrium Mitochondrial Code (transl_table=23)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA * Ter TCA S Ser TAA * Ter TGA * Ter TTG L Leu TCG S Ser TAG * Ter TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu CCG P Pro CAG Q Gln CGG R Arg ATT I Ile i ACT T Thr AAT N Asn AGT S Ser ATC I Ile ACC T Thr AAC N Asn AGC S Ser ATA I Ile ACA T Thr AAA K Lys AGA R Arg ATG M Met i ACG T Thr AAG K Lys AGG R Arg GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val i GCG A Ala GAG E Glu GGG G Gly Click here to change format This code has been created for the mitochondrial genome of the labyrinthulid Thraustochytrium aureum sequenced by The Organelle Genome Megasequencing Program (OGMP). It is the similar to the bacterial code (transl_table 11) but it contains an additional stop codon (TTA) and also has a different set of start codons.Back to top 24. Rhabdopleuridae Mitochondrial Code (transl_table=24)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA * Ter TGA W Trp TTG L Leu i TCG S Ser TAG * Ter TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu i CCG P Pro CAG Q Gln CGG R Arg ATT I Ile ACT T Thr AAT N Asn AGT S Ser ATC I Ile ACC T Thr AAC N Asn AGC S Ser ATA I Ile ACA T Thr AAA K Lys AGA S Ser ATG M Met i ACG T Thr AAG K Lys AGG K Lys GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val i GCG A Ala GAG E Glu GGG G Gly Click here to change format Differences from the Standard Code:
Code 24 Standard
AGA Ser S Arg R
AGG Lys K Arg R
UGA Trp W STOP *
Code 24 has been created for the mitochondrial genome of Rhabdopleura compacta (Pterobranchia). The Pterobranchia are one of the two groups in the Hemichordata which together with the Echinodermata and Chordata form the three major lineages of deuterostomes. AUA translates to isoleucine in Rhabdopleura as it does in the Echinodermata and Enteropneusta while AUA encodes methionine in the Chordata. The assignment of AGG to Lys is not found elsewhere in deuterostome mitochondria but it occurs in some taxa of Arthropoda (Perseke et al. 2011). Code 24 shares with many other mitochondrial codes the reassignment of the UGA STOP to Trp, and AGG and AGA to an amino acid other than Arg. The initiation codons in Rhabdopleura compacta are ATG and GTG (Perseke et al. 2011).Back to top 25. Candidate Division SR1 and Gracilibacteria Code (transl_table=25)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA * Ter TGA G Gly TTG L Leu i TCG S Ser TAG * Ter TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu CCG P Pro CAG Q Gln CGG R Arg ATT I Ile ACT T Thr AAT N Asn AGT S Ser ATC I Ile ACC T Thr AAC N Asn AGC S Ser ATA I Ile ACA T Thr AAA K Lys AGA R Arg ATG M Met i ACG T Thr AAG K Lys AGG R Arg GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val i GCG A Ala GAG E Glu GGG G Gly Click here to change format Differences from the Standard Code:
Code 25 Standard
UGA Gly STOP
Initiation Codons:AUG, GUG, UUG Systematic Range:Candidate Division SR1, Gracilibacteria Comments:Code 25 is used in two groups of (so far) uncultivated Bacteria found in marine and fresh-water environment and in the intestines and oral cavities of mammals among others. The difference to the standard and the bacterial code is that UGA represents an additional glycine codon and does not code for termination (Campbell et al. 2013).Back to top 26. Pachysolen tannophilus Nuclear Code (transl_table=26)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA * Ter TGA * Ter TTG L Leu TCG S Ser TAG * Ter TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG A Ala i CCG P Pro CAG Q Gln CGG R Arg ATT I Ile ACT T Thr AAT N Asn AGT S Ser ATC I Ile ACC T Thr AAC N Asn AGC S Ser ATA I Ile ACA T Thr AAA K Lys AGA R Arg ATG M Met i ACG T Thr AAG K Lys AGG R Arg GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val GCG A Ala GAG E Glu GGG G Gly Click here to change format Differences from the Standard Code:
Code 26 Standard
CUG Ala Leu
Initiation Codons:AUG, GUG, UUG Systematic Range:species in the genera Nakazawaea, Pachysolen and Peterozyma Comments:Code 26 differs from the standard code only the translation of CUG as alanine (as opposed to leucine) (Muhlhausen et al. 2016). It is also similar to code 12 that translates CUG as serine. Code 26 is used by the CUG-Ala clade that include a small number of yeast genera: Nakazawaea, Pachysolen and Peterozyma (Krassowski et al. 2018).Back to top 27. Karyorelict Nuclear Code (transl_table=27)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA Q Gln TGA W Trp TTG L Leu TCG S Ser TAG Q Gln TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu CCG P Pro CAG Q Gln CGG R Arg ATT I Ile ACT T Thr AAT N Asn AGT S Ser ATC I Ile ACC T Thr AAC N Asn AGC S Ser ATA I Ile ACA T Thr AAA K Lys AGA R Arg ATG M Met i ACG T Thr AAG K Lys AGG R Arg GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val GCG A Ala GAG E Glu GGG G Gly Click here to change format Differences from the Standard Code:
Code 27 Standard
UAG Gln STOP
UAA Gln STOP
UGA STOP or Trp STOP
Initiation Codons:AUG Systematic Range:the karyorelictid ciliate Parduczia Comments:Code 27 reassigns the UAG and UAA stops to glutamine while UGA can function as either STOP or tryptophan. Code 27 is used for the karyorelictid ciliate Parduczia sp. (Swart et al. 2016).Back to top 28. Condylostoma Nuclear Code (transl_table=28)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA Q Gln TGA W Trp TTG L Leu TCG S Ser TAG Q Gln TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu CCG P Pro CAG Q Gln CGG R Arg ATT I Ile ACT T Thr AAT N Asn AGT S Ser ATC I Ile ACC T Thr AAC N Asn AGC S Ser ATA I Ile ACA T Thr AAA K Lys AGA R Arg ATG M Met i ACG T Thr AAG K Lys AGG R Arg GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val GCG A Ala GAG E Glu GGG G Gly Click here to change format Differences from the Standard Code:
Code 28 Standard
UAA Gln or STOP STOP
UAG Gln or STOP STOP
UGA Trp or STOP STOP
Initiation Codons: AUGSystematic Range:Condylostoma magnum Comments:Code 28 is used in Condylostoma magnum. The difference to the standard code is that the three stop codons can also be translated as glutamine (UAA, UAG) or tryptophan (UGA), respectively(Swart et al. 2016, Heaphy et al. 2016).Back to top 29. Mesodinium Nuclear Code (transl_table=29)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA Y Tyr TGA * Ter TTG L Leu TCG S Ser TAG Y Tyr TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu CCG P Pro CAG Q Gln CGG R Arg ATT I Ile ACT T Thr AAT N Asn AGT S Ser ATC I Ile ACC T Thr AAC N Asn AGC S Ser ATA I Ile ACA T Thr AAA K Lys AGA R Arg ATG M Met i ACG T Thr AAG K Lys AGG R Arg GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val GCG A Ala GAG E Glu GGG G Gly Click here to change format Differences from the Standard Code:
Code 29 Standard
UAA Tyr STOP
UAG Tyr STOP
Initiation Codons:AUG Systematic Range:the mesodiniid ciliates Mesodinium and Myrionecta Comments:Code 29 is used for the haptorid ciliates Mesodinium and Myrionecta. It differs from the standard code in reassigning the stop codons UAA and UAG to Tyrosine. (Heaphy et al. 2016).Back to top 30. Peritrich Nuclear Code (transl_table=30)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA E Glu TGA * Ter TTG L Leu TCG S Ser TAG E Glu TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu CCG P Pro CAG Q Gln CGG R Arg ATT I Ile ACT T Thr AAT N Asn AGT S Ser ATC I Ile ACC T Thr AAC N Asn AGC S Ser ATA I Ile ACA T Thr AAA K Lys AGA R Arg ATG M Met i ACG T Thr AAG K Lys AGG R Arg GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val GCG A Ala GAG E Glu GGG G Gly Click here to change format Differences from the Standard Code:
Code 30 Standard
UAA Glu STOP
UAG Glu STOP
Initiation Codons:AUG Systematic Range:the peritrich ciliate Carchesium Comments:Code 30 is used in the peritrich ciliate Carchesium. The stop codons UAA and UAG are reassigned to Glutamine. (Sanchez-Silva et al. 2003).Back to top 31. Blastocrithidia Nuclear Code (transl_table=31)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA E Glu TGA W Trp TTG L Leu TCG S Ser TAG E Glu TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu CCG P Pro CAG Q Gln CGG R Arg ATT I Ile ACT T Thr AAT N Asn AGT S Ser ATC I Ile ACC T Thr AAC N Asn AGC S Ser ATA I Ile ACA T Thr AAA K Lys AGA R Arg ATG M Met i ACG T Thr AAG K Lys AGG R Arg GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val GCG A Ala GAG E Glu GGG G Gly Click here to change format Differences from the Standard Code:
Code 31 Standard
UGA Trp STOP
UAG Glu or STOP STOP
UAA Glu or STOP STOP
Initiation Codons:AUG Systematic Range:Blastocrithidia sp. Comments:Code 31 is used for the trypanosome Blastocrithidia sp. UGA encodes trytophan and UAG and UAA encode glutamate and also serve as termination codons. (Zahonova et al. 2016).Back to top 32. Balanophoraceae Plastid Code (transl_table=32)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA * Ter TGA * Ter TTG L Leu i TCG S Ser TAG W Trp TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu i CCG P Pro CAG Q Gln CGG R Arg ATT I Ile i ACT T Thr AAT N Asn AGT S Ser ATC I Ile i ACC T Thr AAC N Asn AGC S Ser ATA I Ile i ACA T Thr AAA K Lys AGA R Arg ATG M Met i ACG T Thr AAG K Lys AGG R Arg GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val i GCG A Ala GAG E Glu GGG G Gly Click here to change format Differences from the Standard Code:
Code 32 Standard
UAG Trp STOP
Systematic Range:Balanophora Comments:Code 32 is used for the highly reduced plastid genome of the holoparasitic plant Balanophora (Su et al. 2018).Back to top 33. Cephalodiscidae Mitochondrial UAA-Tyr Code (transl_table=33)TTT F Phe TCT S Ser TAT Y Tyr TGT C Cys TTC F Phe TCC S Ser TAC Y Tyr TGC C Cys TTA L Leu TCA S Ser TAA Y Tyr TGA W Trp TTG L Leu i TCG S Ser TAG * Ter TGG W Trp CTT L Leu CCT P Pro CAT H His CGT R Arg CTC L Leu CCC P Pro CAC H His CGC R Arg CTA L Leu CCA P Pro CAA Q Gln CGA R Arg CTG L Leu i CCG P Pro CAG Q Gln CGG R Arg ATT I Ile ACT T Thr AAT N Asn AGT S Ser ATC I Ile ACC T Thr AAC N Asn AGC S Ser ATA I Ile ACA T Thr AAA K Lys AGA S Ser ATG M Met i ACG T Thr AAG K Lys AGG K Lys GTT V Val GCT A Ala GAT D Asp GGT G Gly GTC V Val GCC A Ala GAC D Asp GGC G Gly GTA V Val GCA A Ala GAA E Glu GGA G Gly GTG V Val i GCG A Ala GAG E Glu GGG G Gly Click here to change format Differences from the Standard Code:
Code 33 Standard
UAA Tyr STOP
UGA Trp STOP
AGA Ser Arg
AGG Lys Arg
Systematic Range:Cephalodiscidae (Hemichordata) Comments:Code 33 is very similar to the mitochondrial code 24 for the Pterobranchia, which also belong to the Hemichordata, except that it uses UAA for tyrosine rather than as a stop codon (Li Y, Kocot KM, Tassia MG, Cannon JT, Bernt M, Halanych KM. Mitogenomics Reveals a Novel Genetic Code in Hemichordata. Genome Biol Evol. 2019 Jan 1;11(1):29-40.) Back to top
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